We wish to thank the Ocean Drilling Program for sediment samples and especially the core technicians of Leg 169 for providing us minimally disturbed samples as soon as possible after core recovery. Thanks to Barry Cragg for help through the challenges of direct counts in low-abundance samples and to Jody Deming and Steve Summit for providing helpful suggestions on the manuscript. This work was funded by a JOI/USSSP grant to MS and JAB.