This research used samples and data provided by the Ocean Drilling Program (ODP). ODP is sponsored by the U.S. National Science Foundation (NSF) and participating countries under management of Joint Oceanographic Institutions (JOI) Incorporated. Funding for this research was provided by US and international agencies. A.S., H.B., and J.W. thank NWO, the Netherlands Organization for Scientific Research. Funding for the diatom research was provided to CES by the Natural Environment Research Council (NERC)/UK-ODP to CES. CES gratefully acknowledges the Environmental Change Research Centre (University College London, UK).

Reviewers Fabienne Marret, Jens Matthiessen, Mitch Malone, and Lorri Peters are kindly thanked for critical reading and comments. We thank Natasja Welters and Jan van Tongeren (Utrecht) for sample processing. Leonard Bik (Utrecht) is kindly thanked for electronic photography handling and general assistance. The crew of Leg 189 is thanked for general support. This is the Netherlands School of Sedimentary Geology (NSG) contribution number 2003.09.03.